The upper molar cusps is relatively bunoselenomorph

The upper molar cusps is relatively bunoselenomorph

They have a short and you will upright postprotocrista labially established. New preprotocrista, relatively upright, touches a properly-establish parastyle. The metaconule was slightly less which will be large to the M3. Brand new premetaconulecrista in addition to fossae are pretty straight forward instead neocrista. This new rear fossa is actually tilted, specifically to the M3. From upright toward M1, this new postmetaconulecrista becomes sigmoidal on the M3, bypassing the metacone. Brand new labial cusps are in a similar axis that have apparently straight cristae. The fresh paracone labial rib is actually outstanding, but as opposed to prior groove. There’s absolutely no metacone labial rib. The brand new con el fin de-, meso-, and you can metastyles is actually salient. The new lingual cingulum is extremely weak so you’re able to absent for instance the entostyle.

The latest registered structure of your own last straight down premolar is common to own derived base pecoran and you can top pecoran inside having a proper-setup mesolingual conid and posterior cristid , . Furthermore, the current presence of a tooth outlet towards p1 excludes a keen attribution so you can Prodremotherium or Mosaicomeryx . Even when the pearly whites aren’t because the bunoselenodont as with Amphitragulus elegans, this new referred specimens have quite primitive enjoys, such as for instance an incredibly faster metastylid, the lowest crown, a small postentocristid, and though sideways compressed, bulged lingual cuspids to your all the way down molars. Many of these characteristics exclude an affiliation so you can often Dremotherium otherwise Bedenomeryx , . Furthermore, this new very small-size out-of “Amphitragulus” quercyi are an identifying feature into most recent Oligocene ruminants away from Europe , , , .

Dremotherium cf. guthi off Gaimersheim1: step one remaining maxillary bones into tooth row M1-M3 (SNSB-BSPG 1952II4855). out-of Mailhat: 2 kept P4-M2 (MNHN LIM154). Babameryx engesseri away from Gaimersheim1: step three leftover fragmentary mandible having m2 (SNSB-BSPG 1952II4818), (a) lingual, (b) labial, and you can (c) occlusal views. The shape pub is dos cm.

Holotype introduced specimens guidance

Holotype. Jehenne (1987) have designated the latest specimen IPHEP LM.1969.MA.forty as the holotype. This really is a limited right maxillary bone with P3-M3 (, Fig. 2), away from Los angeles Milloque (MP29, France).

Called specimens. Mailhat MNHN LIM154 remaining P4-M2 (that it works, Fig. 6.2); Gaimersheim1 SNSB-BSPG 1952II4855 remaining maxillary having M1-M3 (it really works, Fig. 6.1); SNSB-BSPG 1952II4870 proper m2.


Down long lasting dentition. The reduced molar is extremely defectively managed. But not, so it large ruminant and has a totally selenomorph trigonid. The cristid try elongated as well as the cuspids try large into enamel a bit pleated. New metastylid is well developed.

Top long lasting dentition. New P4 are defectively preserved. Every lingual area try missing. Brand new anterolabial cone and it has a weak labial rib. The fresh new prior stylid are broken. Brand new rear a person is short but establish. No neocrista obstructs new fossa. The newest quadratic upper molars for the occlusal look at incur a proper-arranged metaconule a bit smaller than the protocone. This new cusps are very selenomorph. The fresh new preprotocrista and postmetaconulecrista try labially depending and you will slightly straight. The fresh postprotocrista are a lot of time, highly rounded, and you may labially founded. It can be totally isolated (SNSB-BSPG 1952II4855) otherwise bonded (MNHN LIM154) into the premetaconulecrista. The latter closes from the axis of your own mesostyle. The fresh new posterior basin is has a tiny metaconule fold to your M2, huge to the M3. The brand new paracone, possessing a great paracone rib, as well as the metacone, labially apartment and you can concave, is actually aimed. The brand new mesostyle and you can parastyle are well set up. They setting round small columns. The new mesostyle is huge from M1 so you’re able to M3. The newest entostyle is weakened or missing. There’s none a good lingual cingulum encompassing this new protocone nor a good metastyle creating a properly-arranged cuspid. But not, brief prior and rear cingula exist.


That it ruminant is fairly cutting-edge in-being well-selenodont and having zero lingual cingula during the upper molars, always found in Lophiomeryx, Gelocus, Iberomeryx, Mosaicomeryx, and you will Babameryx , , , [61–63], . In contrast to Bachitherium, that it types does not have a keen anterior groove into very large labial paracone rib , , . The examined fossils much more similar to the advanced stalk Pecora in order to the fresh ancient crown Pecora (instance Amphitragulus, Dremotherium, Bedenomeryx). The fresh enamel is a bit pleated, whereas it’s basically smooth for the Prodremotherium. Furthermore, the dimensions of these pearly whites is much larger than during the Prodremotherium throughout the old Quercy stuff and you may Pech Desse . Tooth be selenomorph compared to “Amphitragulus” and you can Bedenomeryx , , plus the parastyle is not that anteriorly projected than observed into the “Amphitragulus” feningrei , , . Taking a look at the neodiagnosis proposed of the Mennecart , these fossils were all proving top features of Dremotherium. The fresh already outlined species within the European countries is D. feignouxi (type of varieties) and you can D. guthi , , . The latter was a bit smaller compared to the sort kinds , just as the fossils explained right here. D. feignouxi only took place in the Agenian (MN1-MN2; ), while D. guthi existed in the latest Oligocene (MP28-MP30; ). However, from inside the lack of most really-preserved issue, a definitive assignation to your types isn’t feasible.


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